Dinosaur footprints from the Upper Cretaceous of Mongolia

We report new data on 18 dinosaur footprint localities discovered in the Upper Cretaceous of the Gobi Desert of Mongolia, where we have recognized more than 20 000 footprints of dinosaurs. There are at least four types of dinosaur footprints, attributed to theropod, ornithopod, ankylosaurid and sauropod trackmakers. We have also recognized abun dant footprints of un identified trackmakers from each locality. Co existence of footprints and many skeletal remains in the same and/or nearby beds is a remarkable feature of these Mongolian sites. Analyses of dinosaur footprints and associated body fossil remains for each locality reveal that even in the same beds, the ichnofauna differ from the fauna reconstructed on the basis of body fossils of dinosaurs. The results demonstrate that dinosaur faunal assem blages reconstructed from body fossil or footprint evidence solely should be considered very carefully.